Archaeopteryx

George Gaylord Simpson Prize

An Annual Award for a Paper on Evolution and the Fossil Record

Submissions due: Friday, April 3, 2026

The submission deadline has passed.

Yale University graduate students and recent PhD graduates are invited to submit one first-authored paper concerning evolution and the fossil record by the date listed above to the Director’s Office of the Yale Peabody Museum. So long as the paper explicitly addresses the fossil record, the range of questions addressed is open and could include contributions to the philosophy and history of science, theory and methods of phylogenetic inference, biogeography, paleobiology, paleoecology, taphonomy, biostratigraphy, biogeochemical insights into past life, divergence time estimation, biodiversity studies, developmental biology, functional morphology, or conservation biology. Submissions must be either published or in press in a refereed journal. Graduate students in residence in a department at Yale, or past graduates no more than five years after PhD, are eligible. Former winners are not eligible, but papers can be submitted more than once.

The online submission form can be found via link on this web page during active application windows. A copy of the paper must be uploaded as part of the entry.

Winners will be notified via email and will be featured on this web page.

2026 Winner

Jingjun Liu

Jingjun Liu

Evolution of the iodine cycle and the late stabilization of the Earth’s ozone layer

J. Liu, D.S. Hardisty, J.F. Kasting, M. Fakhraee, & N.J. Planavsky, Evolution of the iodine cycle and the late stabilization of the Earth’s ozone layer, Proc. Natl. Acad. Sci. U.S.A. 122 (2) e2412898121, https://doi.org/10.1073/pnas.2412898121 (2025).

Earth’s ozone layer is a crucial prerequisite for the evolution of complex life on land. This research demonstrates that Earth’s evolving marine iodine cycle has fundamentally shaped the stability and abundance of the ozone layer for approximately two billion years following the initial oxygenation of the atmosphere. The resulting elevated flux of UV radiation may have restricted complex life to the ocean and potentially explains the relatively late colonization of multicellular life on land. 

2026 Honorable Mentions

Caleb Gordon

Caleb Gordon

Limb proportions predict aquatic habits and soft-tissue flippers in extinct amniotes

Gordon, C. M., Freisem, L. S., Griffin, C. T., Gauthier, J. A., & Bhullar, B.-A. S. (2025). Limb proportions predict aquatic habits and soft-tissue flippers in extinct amniotes. Current Biology, 35(24), 5982-5998.e7. https://doi.org/10.1016/j.cub.2025.10.068

In this paper, we develop a new machine-learning pipeline, trained on >11,000 original measurements, that can predict whether extinct species were highly or fully aquatic and had soft-tissue flippers with >90% accuracy. Using this pipeline, we reconstruct the evolutionary history of aquatic habits in all major groups of seafaring reptiles, revealing unexpectedly complex patterns and constraints governing their marine invasions. 

Elly Goetz

Elly Goetz

Foraminifera as indicators of species invasion: Ammonia confertitesta in Long Island Sound 

Goetz, E. J., Hull, P. M., Varekamp, J., & Thomas, E. (2025). Foraminifera as indicators of species invasion: Ammonia confertitesta in Long Island Sound. Science Advances, 11(36). https://doi.org/10.1126/sciadv.adv9447 

Determining the origin, introduction, and spread of invasive species is essential for future prevention but is often impossible retroactively. Here, we propose Foraminifera, micro-organisms with calcium carbonate shells, as invasion indicators. Our innovative approach leverages interdisciplinary methods (maritime history, paleontology, ecology) to uncover the timing of and reason(s) for the invasion of the invasive Ammonia confertitesta in Long Island Sound. 

Past Winners

Year Recipient Name
2026 Jingjun Liu
2025

Chase Brownstein

Lisa Freisem 

Alexander Ruebenstahl

2024 Dalton Meyer
2023 Silvina Slagter
2022 Sophie Westacott
2021 Jack Shaw
2020

Michael Hanson

Elizabeth Spriggs

Ross Anderson

2019

Jasmina Wiemann

Christopher Whalen

Daniel Smith-Paredes

2018

Matteo Fabbri

Victoria McCoy

2017

Matt Davis

Nicolas Mongiardino Koch

2016

Simon Darroch

Sarah Federman

Allison Hsaing

2015

Teresa Feo

Alex Dornburg

2014

Rachel Racicot

Stephen Chester

2013 Daniel Field
2012 Una Farrell
2011

Daniel Peppe

April Dinwiddie

2010

Eric Sperling

Jakob Vinther

2009 Faysal Bibi
2008 Jakob Vinther
2007

Walton Green

Brian Moore

Erik A. Sperling

2006

Julia A. Clarke

Ian Miller

2005 Charles D. Bell
2004 Takanobu Tsuihiji
2003

James B. Rossie

Krister T. Smith

2002 Dana Royer
2001 Julia Clarke
2000 Walter Joyce
1998 Daniel Brinkman
1992 Simon Conway Morris
1985 Christine Janis
1984 Karl J. Niklas