George Gaylord Simpson Prize

An Annual Award for a Paper on Evolution and the Fossil Record

Yale University graduate students and recent PhD graduates are invited to submit one first-authored paper concerning evolution and the fossil record by Friday, April 3, 2026 to the Director’s Office of the Yale Peabody Museum. So long as the paper explicitly addresses the fossil record, the range of questions addressed is open and could include contributions to the philosophy and history of science, theory and methods of phylogenetic inference, biogeography, paleobiology, paleoecology, taphonomy, biostratigraphy, biogeochemical insights into past life, divergence time estimation, biodiversity studies, developmental biology, functional morphology, or conservation biology. Submissions must be either published or in press in a refereed journal. Graduate students in residence in a department at Yale, or past graduates no more than five years after PhD, are eligible. Former winners are not eligible, but papers can be submitted more than once.

The online submission form can be found on the Peabody Museum’s website at peabody.yale.edu/george-gaylord-simpson-prize. A copy of the paper must be uploaded as part of the entry.

Winners will be notified via email and will be featured on the Simpson Prize website listed above.

Jingjun Liu

Evolution of the iodine cycle and the late stabilization of the Earth's ozone layer

Earth’s ozone layer is a crucial prerequisite for the evolution of complex life on land. This research demonstrates that Earth's evolving marine iodine cycle has fundamentally shaped the stability and abundance of the ozone layer for approximately two billion years following the initial oxygenation of the atmosphere. The resulting elevated flux of UV radiation may have restricted complex life to the ocean and potentially explains the relatively late colonization of multicellular life on land. 

J. Liu, D.S. Hardisty, J.F. Kasting, M. Fakhraee, & N.J. Planavsky, Evolution of the iodine cycle and the late stabilization of the Earth’s ozone layer, Proc. Natl. Acad. Sci. U.S.A. 122 (2) e2412898121, https://doi.org/10.1073/pnas.2412898121 (2025).

Caleb Gordon

Caleb Gordon

Limb proportions predict aquatic habits and soft-tissue flippers in extinct amniotes

In this paper, we develop a new machine-learning pipeline, trained on >11,000 original measurements, that can predict whether extinct species were highly or fully aquatic and had soft-tissue flippers with >90% accuracy. Using this pipeline, we reconstruct the evolutionary history of aquatic habits in all major groups of seafaring reptiles, revealing unexpectedly complex patterns and constraints governing their marine invasions. 

Gordon, C. M., Freisem, L. S., Griffin, C. T., Gauthier, J. A., & Bhullar, B.-A. S. (2025). Limb proportions predict aquatic habits and soft-tissue flippers in extinct amniotes. Current Biology, 35(24), 5982-5998.e7. https://doi.org/10.1016/j.cub.2025.10.068

Elly Goetz

Elly Goetz

Foraminifera as indicators of species invasion: Ammonia confertitesta in Long Island Sound 

Determining the origin, introduction, and spread of invasive species is essential for future prevention but is often impossible retroactively. Here, we propose Foraminifera, micro-organisms with calcium carbonate shells, as invasion indicators. Our innovative approach leverages interdisciplinary methods (maritime history, paleontology, ecology) to uncover the timing of and reason(s) for the invasion of the invasive Ammonia confertitesta in Long Island Sound. 

Goetz, E. J., Hull, P. M., Varekamp, J., & Thomas, E. (2025). Foraminifera as indicators of species invasion: Ammonia confertitesta in Long Island Sound. Science Advances, 11(36). https://doi.org/10.1126/sciadv.adv9447 

 

Year Recipient Name  
2026 Jingjun Liu  
2025

Chase Brownstein

Lisa Freisem 

Alexander Ruebenstahl

 
2024

Dalton Meyer

 
2023

Silvina Slagter

 
2022

Sophie Westacott

 

2021

Jack Shaw

 

2020

Michael Hanson

Elizabeth Spriggs

Ross Anderson

 

2019

Jasmina Wiemann

Christopher Whalen

Daniel Smith-Paredes

 

2018

Matteo Fabbri

Victoria McCoy

 

2017

Matt Davis

Nicolas Mongiardino Koch

 

2016

Simon Darroch

Sarah Federman

Allison Hsaing

 

2015

Teresa Feo

Alex Dornburg

 

2014

Rachel Racicot

Stephen Chester

 

2013

Daniel Field

 

2012

Una Farrell

 

2011

Daniel Peppe

April Dinwiddie

 

2010

Eric Sperling

Jakob Vinther

 

2009

Faysal Bibi

 

2008

Jakob Vinther

 

2007

Walton Green

Brian Moore

Erik A. Sperling

 

2006

Julia A. Clarke

Ian Miller

 

2005

Charles D. Bell

 

2004

Takanobu Tsuihiji

 

2003

James B. Rossie

Krister T. Smith

 

2002

Dana Royer

 

2001

Julia Clarke

 

2000

Walter Joyce

 

1998

Daniel Brinkman

 

1992

Simon Conway Morris

 

1985

Christine Janis

 

1984

Karl J. Niklas

 


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For questions, please contact:

Sung Yun Senior Administrative Assistant,
Director's Office
+1 203 432 3752 sung.yun@yale.edu